I guess what she meant was that they had a steely, determined personality. I agree with you thought Jayme. I think the characters you mentioned certainly had a determined, take no s**t attitude, but they still were identifiably feminine in appearance. Tifa, for obvious reasons, but Lightning certainly doesn't dress in a masculine way, she shows abit of leg with her outfit, and even Ashe doesn't exactly dress like a nun.
I do sort of see Maybee's point about the main character males being feminine, but only to a point. Famous examples include XIII's Hope Estheim, X's Tidus and VIII's Squall, sort of. Definitely with Hope, be it through his whinyness or tendency to have a slightly high voice, he definitely have some feminine qualities. Tidus and Squall appear masculine, but they have some elements of femininity. However, this isn't the rule. Cloud is definitely quite masculine, Auron is badass, even Snow is the "hero".